Department of Plant Science
Research Interests:

1) Biology of tomato fruit development;

2) Reproductive biology of the tomato wild relatives

3) Plant bioreactor


FAX:  +86-021-34205775  

Address: Department of Plant Science, School of Agriculture and Biology, Shanghai Jiao Tong University




Ph.D. 06/2000
Northeast Agricultural University, School of Horticulture, Harbin, China
Doctor of Philosophy in plant genetics 
Dissertation Title: Study on the genetic resources for the Solanum lycopersicoides [Supervisor: Prof. Jingfu Li]

M.S. 06/1990
Northeast Agricultural University, School of Horticulture, Harbin, China
Masters Degree in plant physiology
Dissertation Title: Dissection of the biomass yield formation for Brassica oleracea during spring [Supervisor: Prof. Shengxuan Li]

Northeast Agricultural University, School of Horticulture, Harbin, China
Bachelors Degree in the horticulture



04/2013~present full time Professor
School of Agriculture and Biology, Fudan-SJTU-Nottingham Plant Biotechnology R&D Center, Shanghai Jiao Tong University, Shanghai, China

11/2011 ~03/2013 Tang Cornell-China Scholar
Department of plant biology, Cornell University, Ithaca, New York, USA

10/2009~10/2011 full time Professor
School of Agriculture and Biology, Fudan-SJTU-Nottingham Plant Biotechnology R&D Center, Shanghai Jiao Tong University, Shanghai, China

04/2009~10/2009 Visiting Professor
Department of Plant Biology, Cornell University, Ithaca, New York, USA

08/2002~12/2008 Associate Professor, full time Professor 
School of Agriculture and Biology, Fudan-SJTU-Nottingham Plant Biotechnology R&D Center, Shanghai Jiao Tong University, Shanghai, China

08/2000-07/2002 Postdoctoral fellow
State Key Laboratory of Genetic Engineering, School of Life Sciences, Fudan University, Shanghai, China

12/1998~07/2000 Associate Research Professor
Harbin Academy of Agricultural Science, Haerbin, China
07/1990-11/1998 Assistant Research Professor
Harbin Academy of Agricultural Science, Haerbin, China    


Teach the courses which including Molecular Biology, Molecular Genetics, Theory of Genetic Engineering and method for undergraduate, graduate and candidate doctor at SJTU's School of Agriculture and Biology, respectively.




  1. 2011 Tang Cornell-China Scholarship Program (11/2011-03/2013)
  2. 2009 State Scholarship Fund. The awardee was selected through a rigid academia evaluation process organized by China Scholarship Council (CSC). This fund sponsors Chinese citizens to pursue study abroad, and certificate presented on March 4, 2009 (Program was completed during 4/2009~10/2009 in Cornell University, USA)
  3. 2009 Teaching Award, “Building and performing of teaching system in research practice for modern agricultural undergraduate students” won second prize in Shanghai city, and the certificate presented in December, 2009
  4. 2008 Teaching Award, “Building of teaching and research practice system for agricultural undergraduate students and performing” won top prize in Shanghai JiaoTong University, and certificate presented on September 15, 2008
  5. 2007 Teaching Award, “Create transgenic lettuce to prevent osteoporosis” won Second prize in Shanghai JiaoTong University for “Challenge Cup” of National Science and Technology College of extra-curricular academic competition works, and certificate presented on November 28, 2007



  1. National Science Foundation of China (NSF), “Dissection of YELLOW-FRUITED TOMAO 3(YFT3) genetic functions for the duration of fruit color development in tomato” (RMB 580,000), Grant No. 32072583, period covered: 01/21-12/2024 (PI)
  2. Shanghai Collaborative Innovation Center of Agri-Seeds(SCCAS), “Creation of novel germplasm and identification of key genes associated with fruit color formation in Solanum lycopersicum”(RMB 500,000) Grant No. ZXWH2150201/010, period covered: 01/21-12/2023 (PI)
  3. Shanghai Tobacco Group Co., Ltd. High Technology Research and Development Program(STG program), “ Creation of the humanized tobacco chassis for plant bioreactor ”(RMB 976,000) , Grant No. WG202102078, period covered: 11/21-12/2022 (PI)
  4. National Science Foundation of China(NSF), “Reveal the molecular mechanism of fruit softening during tomato mature development regulated by YFT1(YELLOW FRUIT TOMATO 1)”(RMB 600,000), Grant No.31872112, period covered: 01/19-12/2022)(PI)
  5. National Science Foundation of China(NSF), “Dissect expression and function of the YFT1(Yellow Fruit Tomato1) gene from ethylene signal transduction to carotenoides biosynthetic pathway for formation of the fruit color in tomato”(RMB 650,000), Grant No.31672158, period covered: 01/17-12/2020)(PI)
  6. National Key Research and Development Program(NKRDP), “Precise identification, innovation and utilization of excellent germplasm resources in the vegetable crops (RMB 420,000), Grant No.2016YFD0100204-28, period covered: 01/16-12/2020)(Co-PI)
  7. Majors program in Science and Technology Commission of Shanghai Municipality, “Dissection of molecular mechanism for fruit color formation in tomato”(RMB 500,000), Grant No.14JC1403400, period covered: 09/2014-08/2017(PI)
  8. National Science Foundation of China(NSF), “Cloning and functional dissection of CD gene related to cuticular cutin in tomato fruit”(RMB330,000), Grant No.31071810, period covered: 01/11-12/2013)(PI)
  9. National high-tech R&D program (863 program), “Develop new techniques of bioreactor to produce therapeutical protein”, Grant No. No.2011AA100607, (RMB1,400,000 to Lingxia, period covered: 08/2011-12/2015)(Co-PI)
  10. National Science Foundation of China(NSF), “Research on the location of fruit specific expression elements and functions of the fused promoter in tomato E8 promotor” (RMB 80,000), Grant No.30871722, period covered:01/2009-12/2009(PI);
  11. National high-tech R&D program(863 program), “Research and development of plant bioreactor to produce fibroblast growth factors and mutant salmon calcitonin”(RMB7,610,000),Grant No.2007AA100503 period covered: 08/2007-10/2010 (PI);
  12. Majors program in Science and Technology Commission of Shanghai Municipality, “Produce thymosin alpha 1 using transgenic plants”(RMB 1,000,000), Grant No.03DZ19310, period covered: 8/2003-12/2006(PI);
  13. National high-tech R&D program (863 program), “Research and development of plant bioreactor to produce mutant salmon calcitonin”(RMB 1,400,000 to Ling Xia), Grant No.2002AA206511, period covered: 01/2002-12/2005(Co-PI).




  1. Chen LL, Li YH, Wang YT, Li WZ, Feng XC and Zhao LX*.Use of high resolution spatiotemporal gene expression data to uncover novel tissue-specific promoters in tomato. MDPI(Multidisciplinary Digital Publishing Institute) Agriculture 2021, 11, 1195.
  2. Zheng CJ, Liu Y, Sun FN, Zhao LX  and Zhang LD. Predicting protein–protein interactions between rice and blast fungus using structure-based approaches. Front. Plant Sci. 12:690124 doi: 10.3389/fpls.2021.690124
  3. Zhao WH, Li YH, Fan SZ, Wen TJ, Wang MH, Zhang LD, Zhao LX. The tomato WRKY32 transcription factor affects ripe fruit color by regulating YFT1, a core component of ethylene signal transduction. Journal of Experimental Botany, 2021,72(12):4269-4282,, 28 May 2021
  4. Zhao WH, Gao L, Li YH, Wang MH, Zhang LD, Zhao LX. Yellow-fruited phenotype is caused by 573 bp insertion at 5’ UTR of YFT1 allele in yft1 mutant tomato. Plant Science, 2020, DOI:
  5. Chen LL,   Li WZ, Li YP,  Feng XC,  Du KY, Wang G, Zhao LX*. Identified trans‑splicing of YELLOW‑FRUITED TOMATO 2 encoding the PHYTOENE SYNTHASE 1 protein alters fruit color by map‑based cloning, functional complementation and RACE. Plant Molecular Biology, 2019,100,647-658,
  6. Ling Li, Weihua Zhao, Xuechao Feng, Lulu Chen, Lida Zhang, and Lingxia Zhao*. Changes in fruit firmness, cell wall composition, and transcriptional profile in the yellow fruit tomato 1 (yft1) mutant. Journal of Agricultural and Food Chemistry (JAFC),2018-12-27, DOI:10.1021/acs.jafc.8b04611;2019,671,463-472
  7. Wang MH, Zhao WH, Gao L, Zhao LX*.Genome-wide profiling of long non-coding RNAs from tomato and a comparison with mRNAs associated with the regulation of fruit ripening. BMC Plant Biology, 2018(18):75,
  8. Liu S, Liu Y, Zhao J, Cai S, Qian H, Zuo K, Zhao L, Zhang L. A computational interactome for prioritizing genes associated with complex agronomic traits in rice (Oryza sativa). Plant Journal,2017,90:177-188. DOI: 10.1111/tpj.13475
  9. Wang YT, Li WZ, Lu C, Fan SZ, Fu CB, Chen MS, Zhao LX*. Evaluation of cultivated tomato germplasm resources. Pakistan Journal Botany, 2017,49(5): 1857-1865
  10. Zhao P F, Pan Q F, Yu W J, Zhao LX*. Dissect style response to pollination using metabolite profiling in selfcompatible and self-incompatible tomato species, Journal of Chromatography B, 2016, 1017: 153-162, 056 DOI information: 10.1016/j.jchromb.2016.01.056
  11. Zhang F Y, Liu SW, Li L, Zuo KJ, Zhao LX, Zhang LD. Genome-wide inference of protein interaction network and its application to the study of crosstalk in Arabidopsis abscisic acid signaling. Plant Physiology2016, DOI:10.1104/pp.16.00057
  12. Gao L, Zhao WH, Qu HO, Wang MH, Zhang LD,Wang QS, Zhao LX*. The yellow fruited tomato 1 (yft1) mutant has altered fruit carotenoid accumulation and reduced ethylene production as a result of a genetic lesion in ETHYLENE INTENSITIVE2.Theor Appl Genet,2016, 129(4):717-728,DOI: 10.1007/s00122-015-2660-4
  13. Zhao PF, Wang M, Zhao LX*. Dissecting stylar responses to self-pollination in wild tomato selfcompatible and self-incompatible species using comparative proteomics. Plant Physiology and Biochemistry, 2016,106 :177-186
  14. Janarthini R, Wang X L, Chen L L, Gao L, Zhao LX*. A tobacco-derived thymosin �4 concatemer promotes cell proliferation and wound healing in mice. BioMed Research International Volume 2016, Article ID 1973413, 8 pages, 
  15. Qu H, Rylosona J SZhao LX.Cytological observation of Solanum pimpinellifolium microspore development. Pakistan Journal of Botany2015, 47(4): 1459-1465. 
  16. Zhao PF, Zhang LD, Zhao LX. Dissection of the style's response to pollination using transcriptome profiling in self-compatible (Solanum pimpinellifolium) and self-incompatible (Solanum chilense) tomato species. BMC plant biology,2015,15:119(2015-04-10) ,DOI 10.1186/s12870-015-0492-7
  17. Gao MF, Peng HZ, Li SS, Wang XL, Gao L, Wang MH, Zhao PF, Zhao LXInsight into flower diversity in Solanum lycopersicum and Solanum chilense using comparative biological approaches. Canadian Journal of Plant Science, 2015, 95(3):467-478.
  18. Gao M, Qu H, Gao L, Chen L, Sebastian R S J, Zhao LDissecting the mechanism of Solanum lycopersicum and Solanum chilense flower colour formation. Plant Biology, 2015,17:1-8, doi:10.1111/plb.12186
  19. Buda G J, Barnes W JFich E APark SYeats TZhao LDomozych D S and Rose J K CAn ABCG transporter is required for cuticular wax deposition and desiccation tolerance in the moss Physcomitrella patensThe Plant Cell, 2013,25:4000-4013
  20. Wang X L, Yang G H, Li SS, Gao MF, Zhao P F, Zhao LX .The Escherichia coli-derived thymosin β 4 concatemer promotes cell proliferation and healing wound in mice. BioMed Research International, 2013, Article ID 241721, 7 pages (Accepted 17 April, in press)
  21. Yeats T H, Martin L B B, Viart H M F, Isaacson T, He Y H, Zhao L X, Matas A J, Buda G J, Domozych D S, Clausen M H, Rose J K C. The identification of cutin synthase: formation of the plant polyester cutin. Nature Chemistry Biology,2012,8:609-611
  22. Cui L J, Peng H Z, Zhang R, Chen Y H, Zhao LX† and Tang KX.The hHscFv-RCR derived from transgenic tobacco acts as a bifunctional molecular complex against Hepatocellular Carcinoma. Biotechnol Appl Biochem.2012,49(1):51-56
  23. Ren W W , Zhao LX, Wang YL, Cui LJ, Tang YL, Sun XF and Tang KX. Overexpression of homogentisate phytyltransferase in lettuce results in increased content of Vitamin E. Afr J Biotechnol 2012,10(64):14046-14051
  24. Ren W W Zhao LX, Zhang LD, Wang YL, Cui LJ, Tang YL, Sun XF and Tang KX. Molecular cloning and characterization of 4-hydroxyphenylpyruvate dioxygenase gene from Lactuca sativa. Journal of Plant Physiology, 2011,168(10): 1076-1083
  25. Ren W, Zhao L, Zhang L, Wang Y, Cui L, Tang Y, Sun X, Tang K. Molecular analysis of a homogentisate phytyltransferase gene from Lactuca sativa L. Mol Biol Rep. 2011,38(3):1813-1819
  26. Cui L J, Chen YH, Shen G A, Zhao L X, Tang KX†. Expression of Bioactive Thymosin Alpha 1 (Tα1) in Marker-free Transgenic Lettuce (Lactuca sativa). Plant Mol Biol Rep,2011, 29:466–472
  27. ZhaoL X, Lu L Y , Zhang L D , Wang A X, Wang N, Liang Z B, Lu X W and Tang  K X. Molecular evolution of the E8 promoter in tomato and some of its relative wild species. Journal of Biosciences, 2009, 34:71–83
  28. Chen Y H, Wang A X, Zhao L X†, Shen G A, Cui L J and Tang K X. Expression of thymosin α1 concatemer in transgenic tomato fruits. Biotechnol Appl Biochem.2009, 2009, 52:303–312 
  29. Chen Y H, Zhao L XShen G A, Cui L J, Ren W W, Zhang HQian H M, Tang K X. Expression and analysis of Thymosin alpha 1 concatemer in Escherichia coliBiotechnol Appl Biochem.2008,49(1):51-56
  30. Zhang HZhao L XChen Y H, Cui L J, Ren W W, Tang K X. Expression of human coagulation factor IX in transgenic tomato. Biotechnol Appl Biochem. 2007,48(2):101-107 
  31. Kai G Y, Miao Z Q, Zhang L, Zhao D, Liao Z H, Sun X F, Zhao L X , Tang K X. Molecular cloning and expression analyses of a new gene encoding 3-hydroxy-3-methylglutaryl-CoA synthase from Taxus media. Biologia Plantarum, 2006, 50 (3): 359–366 
  32. Shen G A, Pang Y Z, Wu W S, Liao Z H, Zhao L X, Sun X F, Tang K X. Cloning and characterizations of a root-specific expressing gene encoding 3-hydroxy-3-methylglutaryl coenzyme A reductase from Ginkgo bilobaMol Biol Rep, 2006, 33:117-127
  33. Zhao L X,Li J F,Chai Y R,Kai G Y,Cao Y F,Sun X F,Tang K X. Investigation on genetic relationship and cross compatibility of S. lycopersiocides and LycopersiconPakistan Journal of Biological Sciences, 2006, 9(6):1160–1168
  34. Shen G A, Pang Y Z, Wu W S, Liu X J, Zhao L X, Sun X F, Tang K X. Isolation and characterization of a putative anthocyanidin reductase gene from Ginkgo bilobaJournal of Plant Physiology, 2006,163: 224-227
  35. Kai GY, Zhao L X, Zhang L, Li Z G, Guo B H, Zhao D L, Sun X F, Miao Z Q, Tang K X. Characterization and expression profile analysis of a new cDNA encoding taxadiene synthase from Taxus media. Joural of Biochemistry and Molecular Biology, 2005, 38(6):668-675
  36. Li Z G*, Zhao L X*, Cui C S,Kai G Y ,Zhang L D,Sun X F, Tang K X. Molecular cloning and characterization of an anti-bolting related gene (BcpFLC3) from Brassica campestris ssp.Pekinensis. Plant science, 2005,168:407–413(* These authors contributed equally to this work)
  37. Shen G A, Pang Y Z, Wu W S, Miao Z Q, Qian H M, Zhao L X, Cao Y F, Sun X F, Tang K X. Molecular cloning, characterization and expression of a novel jasmonate-dependent defensin gene from Ginkgo bilobaJpurnal of Plant Physiology. 2005,162 :1160-1168
  38. Shen G A, Pang Y Z, Wu W S, Deng Z X, Liu X F, Lin J, Zhao L X , Sun X F, Tang K X. Molecular cloning, characterization and expression of a novel Asr gene from Ginkgo bilobaPlant Physiology and Biochemistry, 2005, 43: 836–843
  39. Tang D Q, Qian H M, Zhao L X, Huang D F, Tang K X. Transgenic tobacco plants expressing BoRS1 gene from Brassica oleracea var. acephala show enhanced tolerance water stress. Journal of Biosciences, 2005,30:647-655
  40. Zhao L X , Qiu C X, Li J F, Chai Y R , Kai G.Y, Li Z G, Tang K X. Investigation of disease resistance and cold tolerance of Solanum lycopersicoides for tomato improvement. HortScience, 2005,40 (1):43–46
  41. Kai G Y, Zhao L X, Zheng J G, Zhang L, Zhang L D, Miao Z Q, Li Z G, Sun X F, Tang K X. Isolation and characterization of a new mannose-binding lectin gene from Taxus media. Journal of Bioscience, J. Biosci2004, 29(4): 399–407
  42. Kai G Y, Miao Z Q, Qiu C X, Zhang L, Zhao L X, Li Z G, Xu T F, Zhang L D, Gong Y F, Zhao D L, Liu D H, Sun X F, Tang K X. Molecular cloning and characterization of a taxadienol acetyl transferase cDNA from Taxus mediaPlant science, 2004, 167(4): 759-764
  43. Li Z G*, Zhao L X*, Kai GY,  Yu S W, Cao Y F, Pang Y Z, Sun X F, Tang K X. Cloning and expression analysis of a water stress-induced gene from Brassica oleraceaplant physiology and biochemistry. 2004, 42:789–794(* These authors contributed equally to this work)
  44. Tang D Q, Qian H M, Yu S W, Cao Y F, Liao Z H, Zhao L X, Sun X F, Huang D F, Tang K X.  cDNA cloning and characterization of a new stress-responsive gene BoRS1 from Brassica oleracea var. acephalaPhysiologia Plantarum, 2004, 121: 578-585
  45. Chai Y R, Zhao L X, Liao Z H, Sun X F, Zuo K J, Zhang L, Wang S G, Tang K X. Molecular cloning of potential Verticillium dahliae resistance gene SlVe1 with multi-site polyadenylation from Solanum lycopersicoidesDNA Sequence, 2003,14(5): 375–384
  46. Kai G Y, Zheng J G, Zhang L, Pang Y Z, Liao Z H, Li Z G, Zhao L X, Sun X F, Tang K X. Molecular cloning of a new lectin gene from Z.grandifloraDNA Sequence, 2003,14(4): 335-338
  47. Song W,LI S S, Yang G H, Zhao L X†. Express Recombinant Norovirus P particles in Escherichia coliJournal of Shanghai Jiao Tong University (Agricultural Science),2011,29(2):37-42(Chinese) 
  48. Lu LY, Song W, You X H, Jiang T, Zhao LX. Expression of b ioactive thymosin1 usingmarker free transgenic tobacco. China Biotechnology, 2010, 30 (3): 1-8(Chinese)
  49. Zhang Y, Zhang X P, Zhao LX. Preliminary Research on the Expression of Recombinant Human Proinsulin Using Transgenic Tobacco. Journal of Shanghai Jiao Tong University (Agricultural Science),2010,28(1):14-18, 54(Chinese)
  50. Yang N, Zhao LX, Tang KX. Cloning, expression , and purification of recombinant human-insul in analogue gene in E. coliMedical Journal of Wuhan University, 2010,31(3):281-287 
  51. Cui L J,Lu L Y, Ye J, Pang YZ,Shen GA, Zhao L X† and Tang K X. Development of functional lettuce for preventing and curing osteoporosis. Journal of Shanghai Jiaotong University(Agricultural Science), 2009, 27(5):465-474
  52. Zhao L X, Zhang L, Wei J, Kai G Y, Li Z G, Qian H M, Zhang H, Tang K X. Research on the expression of human coagulation factor IX in tobacco. China Biotechnology, 2006, 26 (3): 11-16(Chinese)
  53. Wang X Y, Pu Y Y, Song J, Wang G F, Qian H M, Lin J, Zhao L X, Tang K X. A preliminary study on the expression of a new type salmon calcitonin gene in transgenic tobacco. Journal of Yangzhou University (Agricultural and Life Science Edition), 2006,27:9-13 (Chinese) († Author for correspondence)
  54. Jin L X, Sun S X, Yu X H, Zhao L X, Li J F, Tang K X. Research on the transgenic tomato using a novel mcgrp gene by Agrobacterium tumefaciens mediated method. Journal of Yangzhou University (Agricultural and Life Science Edition), 2006,27(4):12-16(Chinese)
  55. Zhao L XJin L X, Li Z G, Cao Y F, Qiu C X, Tang D Q, Qian H M, Tang K X. Cloning and DNA sequences analysis of element responsive to ethylene in E8 promoter of the tomato. Acta Horticulturae Sinica, 2004, 31: 204(Chinese)
  56. Zhao L X, Li J F, Kai G Y. Evaluation on the resistance of S olanum lycopersicoides to diseases. Journal of Northeast Agricultural University, 2003, 34(1): 19-23(Chinese)
  57. Zhao L X, Li J F, Xu X Y, Kai GY , Li Z G, Tang K X. Difference on the cold tolerance of Solanum lycopersicoidesActa Horticulturae Sinica, 2003, 30: 416(Chinese)
  58. Zhao L X, Li J F, Tang K X, Xu X Y, Kai G Y. Study on the habits of Solanum lycopersicoides growth and development. Journal of Northeast Agricultural University, 2002, 33(2): 143-146(Chinese)
  59. Zhao L X, Li J F, Tang K X, Xu X Y, Kai G Y. Genetic analysis of Lycopersicon by RAPD markers.Journal of Fudan University (Natural Science Edition),2002,41(2): 222-226 (Chinese)
  60. Zhao L X, Liu S W, Li J F, Xu X Y, Ju W H. Study on optimization of the culture medium for Lycopersicon pollen germination. Journal of Northeast Agricultural University, 2000, 31(2): 147-151(Chinese)

       ​† Corresponding author, * co-first author 




  1. Zhao PF, Peng H Z, Zhao LX. Reproduction strategies of angiosperm plants and Self-incompatibility (In preparation)
  2. Jin LX, Zhao LX*. Research progress, bottlenecks and strategies in the plant bioreactor.China Biotechnology, 2009, 29 (5) : 104-110(Chinese)
  3. Lu LY, Yang N and Zhao L X*. Progress on themolecular biology of the Cladosporium fulvum and its resistance genes in tomato. Acta Horticulturae Sinica, 2009,36:911-922(Chinese)
  4. Meng F J, Huang F L and Zhao L X*. Progress in the Molecular Biology of Solanum peruvianumActa Horticulturae Sinica, 2008,35:1233-1240(Chinese)
  5. Chen Y H, Zhao L X, Cui L X, Tang K X. Progress in transgenic plants producing pharmaceutical proteins. Journal of Agricultural Science and Technology, 2007, 9 (5): 36 – 44(Chinese)
  6. Tang K X, Kai G Y, Zhang L, Pan Z, Zhao L X, Sun X F, Zheng J G. Advances in RACE and its application in plant gene cloning. Journal of Fudan University (Natural Science).2002, 41(6): 704-709(Chinese)
  7. Zhao L X, Li J F, Xu X Y. Advances of the study on Solanum lycopersicoidesActa Horticulturae Sinica, 2000,27:492-496(Chinese)
  8. Zhao L X, Li J.A review: origin, dissemination and classification of LycopersiconCrop Variety and Resource, 1999, 3: 29-31(Chinese)
  9. Zhang W, Yan XF, Zhao LX*. Talking about plant-derived vaccines. Plant Physiology Communications, 2009,45(10):1033-1041(Chinese)

       * Corresponding author




  1. Tomato quality formation, a little story: pursue development of tomato fruit color. Shanhhai Jiao Tong University, May 22-28, 2014(Sino-German Symposium on Metabolomics and Plant Development)
  2. Progress on the plant bioreactor. Northeast Agricultural University, July 21-26, 2011, Harbin, China
  3. Develop a plant oil body system to produce therapeutic recombinant proteins. December 18, 2010, Beijing, China     
  4. estimates the security of plant bioreactor. The 2nd workshop of China national transgenic plant safety, November 27-28, 2007, Beijing, China
  5. Studies on molecular evolution in tomato E8 promoter. The 2nd Annual workshop of China tomato, September 21-24, 2007, Hefei, China
  6. Expresses bioactive thymosinα1 using plant system. April 6-8, 2007, Shanghai, China




  1. Zhao Lingixa, Wang Fu, Li Zhugang, Chai Yourong, Kai Guoyin,Chen Yuhui, Cui Lijie, Meng Fanjuan. “Wild Tomatoes” (in Chinese), Shanghai Jiao Tong University Press(2012-09-01)
  2. Qiao Zongdong, Bai linquan, Zhao lingxia, Miao Xiaoling, Hou Liansheng. “Molecular biology” Shanghai Jiao Tong University Press (2012-05-12). Chapter 9, The molecular biology of mitochonodria and chlorlplasts and Chapter 10, Transposable genetic element.     
  3. Li Jingfu, Wang Aoxue, Xu Xiangyang, Zhao Lingxia, Kang Ligong, Zhang He, Meng Fanjuan. “Research on Tomato Genetics and Breeding (in Chinese)”. Chinese Agricultural Press(2007)




  1. Zhao L X, Cui L J, Zhang R, Song W, Peng H. Designing and synthesis of hHscFv-Rcr and produce hHscFv-Rcr by transgenic tobacco. China Patent Application Serial No. ZL 201010564721.7
  2. Tang K X, Kai G Y, Miao Z Q, Li Z G, Zhao L X. Agglutinin protein coding sequence in Taxus mairei. China Patent No. ZL200410016709.7
  3. Tang K X, Kai G Y, Miao Z Q, Li Z G, Zhao L X. Coding sequence of the taxoid C 13-side-chain-N-benzoyltransferase in Taxus mairei. China Patent No.  ZL200410016769.9
  4. Tang K X, Li Z G, Zhao LX, Sun X F, Kai G Y.Coding sequence of the BcpFLC3 in Brassica pekinensis. China Patent No. ZL03116982.1
  5. Zhao L XLu L Y, Yang N, Zhang Y and Zhang R. Designing and synthesis of the 4×Tαconcatemer gene and develop transgenic tomato to produce thymosin α1. China Patent Application Serial No. 200810041665.1, issued August 14, 2008, Publication Number: 101451137
  6. Zhao L X, Ren W W, Tang K X, Kai G Y, Qian H M, Chen Y H. Designing and synthesis of the hEscFv gene according to the plant codon usage bias. China Patent Application Serial No.200610023236.2, issued January 12, 2006, Publication Number: 1807624
  7. Zhao L X, Cui L J, Tang K X, Kai G Y, Qian H M, Chen Y H. Designing and synthesis of the Rcr gene according to the plant codon usage bias. China Patent Application Serial No. 200610023237.7, issued January 12, 2006, Publication Number: 1807643
  8. Zhao L X, Cui L J, Tang K X, Kai G Y, Qian H M, Chen Y H. Designing and synthesis of the hHscFv gene according to the plant codon usage bias. China Patent Application Serial No.200610023235.8, issued January 12, 2006, Publication Number: 1818063




2009~present, Executive chairman for ‎Young Scientists forum in Field Horticulture 

2008~present, China Society of tomato

2007~present, China Society of Plant Physiologists

2005~present, Chinese Society of Biotechnology

2000~present, China Society of botany

1998~present, China Society of Horticulture




  1. Regular reviewer of competitive grants National Natural Science Foundation of China
  2. Regular reviewer for Natural Science Foundation of Zhejiang Province, and Jiangsu Province
  3. Peer reviewer for Molecular Biology ReportsPhysiologia PlantarumJournal of the American Society for Horticultural ScienceJournal of Plant Growth RegulationBiotechnology and Applied Biochemistry, Chemistry of life, Agricultural Science in China

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